Big, Busy and Complex: Digg.com's Web Infrastructure
Further to the note about TurboTax (below), ComputerWord has an article that gives a good insight into the commitment, investment and skills needed to operate the infrastructure of a very Large Scale website.
Digg.com is a peer review community whereby site subscribers submit and vote on interesting web articles. In order to cope the huge demands of its 1.2 MILLION users, Digg now runs over 100 separate servers. This includes 30 webservers, 20 database servers, as well as several search servers (the rest are used for backup).
The article reveals how important decisions about infrastructure management are for the performance of a large and busy website.
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